While printing, the "Tuning Adjustment" meters allow you to adjust the speed of the extruder as well as the amount of filament being extruded. This can be helpful if you realize mid-print that the extruder is moving too fast for the filament to stick or if not enough filament is being extruded. These are the only settings you can adjust during a print; otherwise, you have to cancel the print, adjust the settings and start over.
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